Phylogenetic support Arrhenia consistently appears as a paraphyletic grade in all analyses, and the same is true for tribe Arrhenieae. Species included Type

species: Arrhenia auriscalpium. Species included based on molecular phylogeny are A. chlorocyanea (Pat.) this website Redhead et al., Lutzoni, Moncalvo & Vilgalys, A. epichysium (Pers. : Fr.) Redhead et al., A. griseopallida (Desm.) Watling, A. lobata (Pers.) Kühner & Lamoure ex Redhead, A. obscurata (D.A. Reid) Redhead et al., A. philonotis (Lasch) Redhead et al., A. sphagnicola (Berk.) Redhead et al. and A. velutipes (P.D. Orton) Redhead et al. Species included in Arrhenia based on morphology in Redhead et al. (2002) are A. acerosa (Fr.) Kühner, A. alnetora (Singer) Redhead, A. australis (Clel.) Grgurinovic, A. andina (Corner) Redhead et al., A. antarctica (Singer) Redhead et AMN-107 al., A. baeospora (Singer) Redhead et al., A. chilensis (Mont.) Redhead et al., A. elegans (Pers.) Redhead et al., A. fissa (Leyss.) Redhead, A. hohensis (A.H. Sm.) Redhead et al., A. lundellii (Pilát) Redhead et al., A. obatra (J. Favre) Redhead et al., A. obscurata (D. A. Reid) Redhead et al., A. omnivora (Agerer) Redhead et al., A. onisca (Fr.:Fr.) Redhead et al., A. parvivelutina (Clémençon & Irlet) Redhead et al., A. pauxilla

(Clémençon) selleck compound Redhead et al., A. peltigerina (Peck) Redhead et al., A. pubescentipes (H.E. Bigelow) Redhead et al., A. rainierensis (H.E. Bigelow) Redhead et al., A. retiruga Redhead, A. rickenii (Hora) Watling, A. rigidipes (Lamoure) Redhead et al., A. salina (Høil.) Bon & Courtec., A. spathulata (Fr.) Redhead, A. rustica (Fr.) Redhead et Farnesyltransferase al., A. sphaerospora (Lamoure) Redhead et al., A. stercoraria (Barrasa, Esteve-Rav. & Sánchez Nieto) Redhead et al., A. subglobispora (G. Moreno, Heykoop & E. Horak) Redhead et al., A. subobscura (Singer) Redhead et al., A. subumbratilis Redhead et al., A. trigonospora (Lamoure) Redhead et al., A. umbratilis

(Fr.:Fr.) Redhead et al., A. viridimammata (Pilát) Redhead et al. and A. volkertii (Murrill) Redhead et al. Comments Omphalinoid Arrhenia species were once classified in Omphalina (type species, O. pyxidata), a genus that is also bryophilous, but Arrhenia are gray-brown throughout while Omphalina have a reddish brown surface and colorless context (Redhead et al. 2002). Arrhenia was erected for species with drooping or pendant basidiomata with cantharelloid (wrinkled) hymenia (Corner 1966, Høiland 1976; Pilát and Nannfeldt 1954), but later expanded to include species with pleurotoid basidiomata, such as Leptoglossum and Phaeotellus, and omphalinoid basidiomata (Redhead et al. 2002). Because Arrhenia includes reduced species (e.g., A. auriscalpium, the type of Arrhenia, and A. lobata, the type of Dictyolus Quél.) as well as omphalinoid species, some are not readily distinguishable from other genera in the subfamily based on macromorphology.